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These 8 cells then divide and produce the millions of cells that make up a cat. Different cells inactivate different copies of the X chromosome, which means some cells express alleles for orange and white and other cells express black and white alleles. X chromosome inactivation occurs at the 8 cell stage of the embryo. Xist and Tsix have mutually exclusive expression, which ensures that only one X chromosome is inactivated in each cell. (For further details of the process, check out my post on Nessa Carey's book Junk DNA.) In short, a long non-coding RNA known as Xist (X-inactive specific transcript) turns off one copy of the X chromosome in each cell on the opposite strand of DNA, a gene called Tsix is found. The specifics of how this happens is actually pretty amazing. In organisms like the fruit fly, the XY males just double the expression of the genes on the X chromosome. In mammals, this involves silencing one of the X chromosomes in each cell with the XX genotype. Organisms have different ways to level the compensate for these differences. If an XX individual expressed all the genes on both X chromosomes, they would have a huge imbalance in protein expression in comparison with someone with XY. In contrast, the X chromosome contains lots of protein-encoding genes. The Y chromosome is rather small and does not encode many genes. Somatic chromosomes come in even pairs, but the sex chromosomes are not created equal. The patterns observed in calico and tortoiseshell cats can be explained by a phenomenon called X chromosome inactivation. If fur color followed the rules of Mendelian genetics, you would expect cats to be only black and white or orange and white, not two or three colors at once. The genetics of the tortoiseshell/calico cat The gene for white spotting (S) has been linked to the KIT locus more recent publications have shown that a retroviral insertion in the KIT oncogene caused white spotting, with a full insertion leading to the recessive all white mutation. However, recessive versions of the MC1R allele create orange/amber fur color in Norwegian forest cats. A recent study of the Syrian hamster, which shows sex-linked inheritance of yellow fur color, described the sex-linked yellow gene ( Sly) as being independent of MC1R function and not likely homologous to the Orange gene in cats. Interestingly, the MC1R gene, which has been linked to red hair in humans and other mammals, is not involved in the coloring of calico cats because it isn't on the X chromosome. The Orange gene has not yet been identified, but its position on the X chromosome has been mapped. There are rare cases of male cats with Klinefelter syndrome (XXY) that can have these fur color variants.Īgouti (gene name ASIP) encodes a protein that inactivates the melanocortin receptor and thus controls the distribution and amount of pigment cells (melanocytes) in the hair. Because the Orange gene is on the X chromosome, most torties and calico cats are female. Based on the combined expression of these genes, some cats are tortoiseshell, a mix of black and orange and some cats are calico, a mix of black, orange, and white. For this discussion, I will focus only on the gene loci that create the calico pattern: agouti (A), orange (O), and spotted (S). In other mammals, white fur color alone has been linked to at least six genes ( MITF, EDN3, EDNRB, PAX3, SOX10, and SNAI2). There are many different genes that create the spectrum of coat colors and patterns found in domestic cats. The genetics of cat fur color (like the genetics of human hair color) are rather complex. While researching the post, I found myself in another internet rabbit hole and I wanted to share some of the things I learned about cat genetics because, it turns out, calico cats are an excellent lesson in genetics. Nettie Stevens is known for her discovery of the X and Y chromosomes as the basis for sex determination. In an earlier post, I detailed the life and work of Nettie Maria Stevens, the namesake of our new calico cat. These patched tabbies are often called a tortie-tabby, a torbie or, with large white areas, a caliby. Occasionally tabby patterns of black and brown (eumelanistic) and red (phaeomelanistic) colours are also seen. Dilution genes may modify the colouring, lightening the fur to a mix of cream and blue, lilac or fawn the markings on tortoiseshell cats are usually asymmetrical. Typically, the more white that a cat has, the more solid the patches of colour. The size of the patches can vary from a fine speckled pattern to large areas of colour. Tortoiseshell cats have particoloured coats with patches of various shades of red, grey, and black, and sometimes white. Patterns Cat with a blue ("dilute") tortoiseshell coat A tortoiseshell with characteristic "split-face" pattern This pattern is especially preferred in the Japanese Bobtail breed, and exists in the Cornish Rex group. Tortoiseshell markings appear in many different breeds, as well as in non-purebred domestic cats. Those that are predominantly white with tortoiseshell patches are described as tricolour, tortoiseshell-and-white, or calico (in Canada and the United States). "Tortoiseshell" is typically reserved for multicoloured cats with relatively small or no white markings. Tortoiseshell cats with the tabby pattern as one of their colours are sometimes referred to as torbies or torbie cats. The colours are often described as red and black, but the "red" patches can instead be orange, yellow, or cream, and the "black" can instead be chocolate, grey, tabby, or blue. Tortoiseshell cats combine two colours other than white, either closely mixed or in larger patches. Male tortoiseshells are rare and are usually sterile. Like tortoiseshell-and-white or calico cats, tortoiseshell cats are almost exclusively female. Tortoiseshell is a cat coat colouring named for its similarity to tortoiseshell pattern. Some of them are best magazine themes, some of them are blog themes and some of them are multipurpose as well. Best Free WordPress themes for 2023Īll the below-listed themes are popular in their own niche. Some theme has fewer active installs, but still, they are awesome. This collection is made based on active installs, user feedback, ratings, and support availability. In this blog post, we are going to share some best the best free WordPress themes and templates for your WordPress website. However, the free version may lack some useful features and dedicated support on time. 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Urn:lcp:tooheysmedicinef0000tooh_h0a9:epub:ac93cdcb-f0dc-44c9-b654-33a4876a1860 Foldoutcount 0 Identifier tooheysmedicinef0000tooh_h0a9 Identifier-ark ark:/13960/t22c9498q Invoice 1652 Isbn 0443030766 Lccn 85016680 Ocr tesseract 4.1.1 Ocr_detected_lang en Ocr_detected_lang_conf 1.0000 Ocr_detected_script Latin Ocr_detected_script_conf 0.9889 Ocr_module_version 0.0.11 Ocr_parameters -l eng Old_pallet IA-NS-2000215 Openlibrary_edition In spite of recent changes in nurse education, nurses still need to know about medicine to understand their work with patients. Access-restricted-item true Addeddate 22:11:00 Associated-names Bloom, Arnold Bloom, Stephen Robert Boxid IA40056411 Camera Sony Alpha-A6300 (Control) Collection_set printdisabled External-identifier Description Creators/Contributors Contributor Bloom, Stephen Robert. Made with deltamethrin, this insecticide is ideal for use around the home to control ants, bed bugs, cockroaches, crickets, fleas, firebrats, silverfish, termites, and more. 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